If Darwin had been aware of genes, and of their typical mode of transmission to subsequent generations, with which statement would he most likely have been in agreement?

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Darwin's 4 theories of natural selection are: The four propositions underlying Darwin's theory of evolution through natural selection are:

(1) more individuals are produced than can survive; (2) there is therefore a struggle for existence; (3) individuals within a species show variation; and (4) offspring tend to inherit their parents' characters.


If Darwin had been aware of genes, and of their typical mode of transmission to subsequent generations, with which statement would he most likely have been in agreement?