An ecosystem is joined by thousands of species, if a problem occurs causing a species to become extinct, it may not affect the proper functioning of the ecosystem initially, but as more and more species are removed, the ecosystem becomes dangerously weak over a period of time. Which hypothesis gave this sentence?

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An ecosystem is joined by thousands of species, if a problem occurs causing a species to become extinct, it may not affect the proper functioning of the ecosystem initially, but as more and more species are removed, the ecosystem becomes dangerously weak over a period of time. Which hypothesis gave this sentence?