Name the basic structural unit of chromatin, which is described by R.Kornberg?

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Eukaryotic chromosomes consist of DNA tightly wound around clusters of histone proteins. In general, eukaryotic cells contain a lot more genetic material than prokaryotic cells. For example, each human cell has around 2m, or 3 billion base pairs, of DNA that must be compacted to fit within the nucleus.
 

Chromatin is a complex of DNA and proteins that forms chromosomes within the nucleus of eukaryotic cells. Nuclear DNA does not appear in free linear strands; it is highly condensed and wrapped around nuclear proteins in order to fit inside the nucleus.


Name the basic structural unit of chromatin, which is described by R.Kornberg?